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mx id: 1627 | OBO id: HAO:0000390 | URI: http://purl.obolibrary.org/obo/HAO_0000390
Defined (both definition and relationships must be met)
The carina that is transverse and posteriorly directed laterally and is located on the abdominal tergite.
written by: Miko, I., M. Bertone, M. Yoder, A. Deans, S. Katja, N. Johnson, S. Brady, P. Ward, F. Ronquist, E. Talamas, L. Krogmann, L. Vilhelmsen, C. Rasmussen, G. Broad, A. Lucky, and B. G. 2010. HAO/Biosync working meeting II..

Relationships / properties:
The gradulus is a carina.
The gradulus is part of the cuticle. The gradulus is part of the metasoma. The gradulus is part of the sclerite.

Label usage (sensu)
gradulus by Deans, A. R. 2009. HAO curator..
gradulus by Bohart, R. M., and A. S. Menke. 1976. Sphecid Wasps of the World, A Generic Revision. University of California Press, Berkeley, CA 695 pp.
Comments and tags
alternative definition added over 10 years ago by Andy Deans
a line that runs from side to side on metasomal segments that is formed by the area between two regions of different height (NOT sure of origin; reference is best guess)
  Reference: Michener, C. D. 2000. The Bees of the World. Johns Hopkins University Press, Baltimore, MD, USA 913 pp.
  Cross reference: None provided.

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